Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure

In flowers of Nymphaea and Victoria, carpellary appendages are regarded as structures related to pollination by deceit of night-blooming species. In this study, the anatomy, histochemistry and ultrastructure of carpellary appendages were analysed to investigate their possible role in the production...

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Autores principales: Zini, Lucía Melisa, Galati, Beatriz Gloria, Gotelli, Marina Maria, Zarlavsky, Gabriela Elena, Ferrucci, María Silvia
Formato: Artículo
Lenguaje:Inglés
Publicado: Wiley Blackwell Publishing, Inc 2025
Acceso en línea:http://repositorio.unne.edu.ar/handle/123456789/59950
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spelling I48-R184-123456789-599502025-12-15T15:19:44Z Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure Zini, Lucía Melisa Galati, Beatriz Gloria Gotelli, Marina Maria Zarlavsky, Gabriela Elena Ferrucci, María Silvia In flowers of Nymphaea and Victoria, carpellary appendages are regarded as structures related to pollination by deceit of night-blooming species. In this study, the anatomy, histochemistry and ultrastructure of carpellary appendages were analysed to investigate their possible role in the production of volatile compounds in nocturnal species Nymphaea amazonum, N. gardneriana, N. prolifera (Nymphaea subgenus Hydrocallis) and Victoria cruziana, and in diurnal species N. caerulea (Nymphaea subgenus Brachyceras). Carpellary appendages were studied using light microscopy and scanning and transmission electron microscopy from pre-anthesis to the second day of anthesis. Anatomical and ultrastructural features are characteristic of osmophores. In all species, the most frequent components in secretory cells are amyloplasts, lipid bodies, mitochondria, rough endoplasmic reticulum and elaioplasts. The epidermis and multilayered parenchyma accumulate abundant starch grains and lipophilic substances, both of which vanish during anthesis. Amorphous substances are deposited between the plasmalemma and the outer cell wall of epidermal cells, and are then released by cuticular diffusion. Odour production in carpellary appendages might be an ancient role of primary importance both in diurnal and nocturnal species that are pollinated by deceit. Olfactory and visual cues of small carpellary appendages in Nymphaea subgenus Brachyceras correspond to bee pollination, and large carpellary appendages in subgenus Hydrocallis and Victoria represent parallel functional specializations of the flowers to the attraction and reward for exclusive beetle pollination. 2025-12-15T14:21:32Z 2025-12-15T14:21:32Z 2019 Artículo Zini, Lucía Melisa, et al., 2019. Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure. Reino Unido: Wiley Blackwell Publishing, Inc, vol. 191, no. 4, p. 421-439. E-ISSN 1095-8339. 0024-4074 http://repositorio.unne.edu.ar/handle/123456789/59950 eng https://academic.oup.com/botlinnean/search-results?page=1&q=Carpellary%20appendages%20in%20Nymphaea%20and%20Victoria%20(Nymphaeaceae)%3A%20evidence%20of%20their%20role%20as%20osmophores%20based%20on%20morphology%2C%20anatomy%20and%20ultrastructure&fl_SiteID=5596&SearchSourceType=1&allJournals=1 closedAccess http://creativecommons.org/licenses/by-nc-nd/2.5/ar/ application/pdf p. 421-439 application/pdf Wiley Blackwell Publishing, Inc Botanical Journal of the Linnean Society, 2019, vol. 191, no. 4, p. 421-439.
institution Universidad Nacional del Nordeste
institution_str I-48
repository_str R-184
collection RIUNNE - Repositorio Institucional de la Universidad Nacional del Nordeste (UNNE)
language Inglés
description In flowers of Nymphaea and Victoria, carpellary appendages are regarded as structures related to pollination by deceit of night-blooming species. In this study, the anatomy, histochemistry and ultrastructure of carpellary appendages were analysed to investigate their possible role in the production of volatile compounds in nocturnal species Nymphaea amazonum, N. gardneriana, N. prolifera (Nymphaea subgenus Hydrocallis) and Victoria cruziana, and in diurnal species N. caerulea (Nymphaea subgenus Brachyceras). Carpellary appendages were studied using light microscopy and scanning and transmission electron microscopy from pre-anthesis to the second day of anthesis. Anatomical and ultrastructural features are characteristic of osmophores. In all species, the most frequent components in secretory cells are amyloplasts, lipid bodies, mitochondria, rough endoplasmic reticulum and elaioplasts. The epidermis and multilayered parenchyma accumulate abundant starch grains and lipophilic substances, both of which vanish during anthesis. Amorphous substances are deposited between the plasmalemma and the outer cell wall of epidermal cells, and are then released by cuticular diffusion. Odour production in carpellary appendages might be an ancient role of primary importance both in diurnal and nocturnal species that are pollinated by deceit. Olfactory and visual cues of small carpellary appendages in Nymphaea subgenus Brachyceras correspond to bee pollination, and large carpellary appendages in subgenus Hydrocallis and Victoria represent parallel functional specializations of the flowers to the attraction and reward for exclusive beetle pollination.
format Artículo
author Zini, Lucía Melisa
Galati, Beatriz Gloria
Gotelli, Marina Maria
Zarlavsky, Gabriela Elena
Ferrucci, María Silvia
spellingShingle Zini, Lucía Melisa
Galati, Beatriz Gloria
Gotelli, Marina Maria
Zarlavsky, Gabriela Elena
Ferrucci, María Silvia
Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
author_facet Zini, Lucía Melisa
Galati, Beatriz Gloria
Gotelli, Marina Maria
Zarlavsky, Gabriela Elena
Ferrucci, María Silvia
author_sort Zini, Lucía Melisa
title Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
title_short Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
title_full Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
title_fullStr Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
title_full_unstemmed Carpellary appendages in Nymphaea and Victoria (Nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
title_sort carpellary appendages in nymphaea and victoria (nymphaeaceae) : evidence of their role as osmophores based on morphology, anatomy and ultrastructure
publisher Wiley Blackwell Publishing, Inc
publishDate 2025
url http://repositorio.unne.edu.ar/handle/123456789/59950
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