Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea

The genus Larrea has an amphitropical distribution in North and South American deserts, and its phylogeny remains unresolved. This genus is conspicuous and specious within the Monte Desert, the largest, although understudied, southern South American dryland. Larrea presents an interesting case for...

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Autores principales: Quiroga, María Paula, Castello, Lucia V., Tadey, Mariana, Márquez, Sebastián, Premoli, Andrea C., Souto, Cintia P.
Formato: Articulo article acceptedVersion
Lenguaje:Inglés
Publicado: Adrian Brennan 2025
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Acceso en línea:https://rdi.uncoma.edu.ar/handle/uncomaid/18835
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id I22-R178-uncomaid-18835
record_format dspace
institution Universidad Nacional del Comahue
institution_str I-22
repository_str R-178
collection Repositorio Institucional UNCo
language Inglés
topic Chloroplast sequences
Creosote bush
Cryptic hybridization
Hybrid swarm
Monte Desert
North and South American drylands
Nuclear sequences
Ciencias de la Tierra y Medio Ambiente
spellingShingle Chloroplast sequences
Creosote bush
Cryptic hybridization
Hybrid swarm
Monte Desert
North and South American drylands
Nuclear sequences
Ciencias de la Tierra y Medio Ambiente
Quiroga, María Paula
Castello, Lucia V.
Tadey, Mariana
Márquez, Sebastián
Premoli, Andrea C.
Souto, Cintia P.
Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
topic_facet Chloroplast sequences
Creosote bush
Cryptic hybridization
Hybrid swarm
Monte Desert
North and South American drylands
Nuclear sequences
Ciencias de la Tierra y Medio Ambiente
description The genus Larrea has an amphitropical distribution in North and South American deserts, and its phylogeny remains unresolved. This genus is conspicuous and specious within the Monte Desert, the largest, although understudied, southern South American dryland. Larrea presents an interesting case for phylogenetic studies due to its paternally inherited chloroplasts, its species hybridize in nature, and although nominal species are morphologically distinct, hybrids might be cryptic. We analysed ITS2 nuclear (nDNA) and rbcL chloroplast (cpDNA) sequences of the bifoliolate section, Bifolium, including L. tridentata (Lt) from North America, and its South American congeners: L. cuneifolia (Lc) and L. divaricata (Ld), and sequences of the multifoliolate Larrea section: L. ameghinoi (La), L. nitida (Ln), and a morphological hybrid swarm. We aligned and analysed sequences from 111 individuals collected at 31 populations sampled along the range of each species. The nDNA revealed 56 haplotypes, and median-joining and maximum likelihood reconstructions provided clear separation among species and suggested hybridization between Lc-Ld. The nuclear phylogeny showed that the section Larrea diverged earlier than Bifolium, within which Lc diverged first, meanwhile, consistent with previous studies, Lt forms a monophyletic group sister to Ld. Comparatively, cpDNA was less variable, with only six haplotypes shared between Ln-Lc and Ln-La, and rarely between Ld-Lc. Our results emphasize the significance of separately considering nuclear and plastid evolutionary signals when reconstructing unresolved relationships. While nuclear markers clarified phylogenetic relationships and cryptic hybridization among Larrea species, the chloroplast revealed the retention of widespread ancient polymorphisms, which were conserved in populations of distinct species. Each marker provided insights into particular evolutionary patterns, highlighting that genetic variation may be more influenced by hybridization and mode of chloroplast inheritance than previously recognized.
format Articulo
article
acceptedVersion
author Quiroga, María Paula
Castello, Lucia V.
Tadey, Mariana
Márquez, Sebastián
Premoli, Andrea C.
Souto, Cintia P.
author_facet Quiroga, María Paula
Castello, Lucia V.
Tadey, Mariana
Márquez, Sebastián
Premoli, Andrea C.
Souto, Cintia P.
author_sort Quiroga, María Paula
title Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
title_short Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
title_full Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
title_fullStr Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
title_full_unstemmed Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea
title_sort everything everywhere all at once? disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus larrea
publisher Adrian Brennan
publishDate 2025
url https://rdi.uncoma.edu.ar/handle/uncomaid/18835
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spelling I22-R178-uncomaid-188352025-07-22T17:53:04Z Everything everywhere all at once? Disentangling the long-lasting riddle of phylogenetic relationships and cryptic hybridization in the amphitropical genus Larrea Quiroga, María Paula Castello, Lucia V. Tadey, Mariana Márquez, Sebastián Premoli, Andrea C. Souto, Cintia P. Chloroplast sequences Creosote bush Cryptic hybridization Hybrid swarm Monte Desert North and South American drylands Nuclear sequences Ciencias de la Tierra y Medio Ambiente The genus Larrea has an amphitropical distribution in North and South American deserts, and its phylogeny remains unresolved. This genus is conspicuous and specious within the Monte Desert, the largest, although understudied, southern South American dryland. Larrea presents an interesting case for phylogenetic studies due to its paternally inherited chloroplasts, its species hybridize in nature, and although nominal species are morphologically distinct, hybrids might be cryptic. We analysed ITS2 nuclear (nDNA) and rbcL chloroplast (cpDNA) sequences of the bifoliolate section, Bifolium, including L. tridentata (Lt) from North America, and its South American congeners: L. cuneifolia (Lc) and L. divaricata (Ld), and sequences of the multifoliolate Larrea section: L. ameghinoi (La), L. nitida (Ln), and a morphological hybrid swarm. We aligned and analysed sequences from 111 individuals collected at 31 populations sampled along the range of each species. The nDNA revealed 56 haplotypes, and median-joining and maximum likelihood reconstructions provided clear separation among species and suggested hybridization between Lc-Ld. The nuclear phylogeny showed that the section Larrea diverged earlier than Bifolium, within which Lc diverged first, meanwhile, consistent with previous studies, Lt forms a monophyletic group sister to Ld. Comparatively, cpDNA was less variable, with only six haplotypes shared between Ln-Lc and Ln-La, and rarely between Ld-Lc. Our results emphasize the significance of separately considering nuclear and plastid evolutionary signals when reconstructing unresolved relationships. While nuclear markers clarified phylogenetic relationships and cryptic hybridization among Larrea species, the chloroplast revealed the retention of widespread ancient polymorphisms, which were conserved in populations of distinct species. Each marker provided insights into particular evolutionary patterns, highlighting that genetic variation may be more influenced by hybridization and mode of chloroplast inheritance than previously recognized. Fil: Quiroga, María Paula. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Quiroga, María Paula. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. Fil: Castello, Lucia V. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Castello, Lucia V. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. Fil: Tadey, Mariana. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Tadey, Mariana. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. Fil: Márquez, Sebastián. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Márquez, Sebastián. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. Fil: Premoli, Andrea C. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Premoli, Andrea C. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. Fil: Souto, Cintia P. Universidad Nacional del Comahue. Centro Regional Universitario Bariloche; Argentina. Fil: Souto, Cintia P. Consejo Nacional de Investigaciones Científicas y Técnicas. Instituto de Investigaciones en Biodiversidad y Medioambiente; Argentina. 2025 2025-07-04T15:44:37Z 2025-07-04T15:44:37Z Articulo article acceptedVersion https://rdi.uncoma.edu.ar/handle/uncomaid/18835 eng https://doi.org/10.1093/aobpla/plaf024 Atribución-NoComercial-CompartirIgual 4.0 https://creativecommons.org/licenses/by-nc-sa/4.0/ application/pdf application/pdf Adrian Brennan Oxford Academic AoB PLANTS, Volume 17, Issue 3, June 2025, plaf024