Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling

Each heartbeat is followed by a refractory period. Recovery from refractoriness is known as Ca²⁺ release restitution (CRR), and its alterations are potential triggers of Ca²⁺ arrhythmias. Although the control of CRR has been associated with SR Ca²⁺ load and RYR2 Ca²⁺ sensitivity, the relative role o...

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Autores principales: Cely Ortiz, Diana Catalina Alejandra, Felice, Juan Ignacio, Diaz Zegarra, Leandro Agustín, Valverde, Carlos Alfredo, Federico, Marilén, Palomeque, Julieta, Wehrens, Xander H. T., Kranias, Evangelia G., Aiello, Ernesto Alejandro, Lascano, Elena Catalina, Negroni, Jorge A., Mattiazzi, Alicia Ramona
Formato: Articulo
Lenguaje:Inglés
Publicado: 2020
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Acceso en línea:http://sedici.unlp.edu.ar/handle/10915/124703
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id I19-R120-10915-124703
record_format dspace
institution Universidad Nacional de La Plata
institution_str I-19
repository_str R-120
collection SEDICI (UNLP)
language Inglés
topic Medicina
Cellular Physiology
Computational biology
Intercellular Signaling
Molecular Physiology
spellingShingle Medicina
Cellular Physiology
Computational biology
Intercellular Signaling
Molecular Physiology
Cely Ortiz, Diana Catalina Alejandra
Felice, Juan Ignacio
Diaz Zegarra, Leandro Agustín
Valverde, Carlos Alfredo
Federico, Marilén
Palomeque, Julieta
Wehrens, Xander H. T.
Kranias, Evangelia G.
Aiello, Ernesto Alejandro
Lascano, Elena Catalina
Negroni, Jorge A.
Mattiazzi, Alicia Ramona
Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
topic_facet Medicina
Cellular Physiology
Computational biology
Intercellular Signaling
Molecular Physiology
description Each heartbeat is followed by a refractory period. Recovery from refractoriness is known as Ca²⁺ release restitution (CRR), and its alterations are potential triggers of Ca²⁺ arrhythmias. Although the control of CRR has been associated with SR Ca²⁺ load and RYR2 Ca²⁺ sensitivity, the relative role of some of the determinants of CRR remains largely undefined. An intriguing point, difficult to dissect and previously neglected, is the possible independent effect of SR Ca²⁺ content versus the velocity of SR Ca²⁺ refilling on CRR. To assess these interrogations, we used isolated myocytes with phospholamban (PLN) ablation (PLNKO), knock-in mice with pseudoconstitutive CaMKII phosphorylation of RYR2 S2814 (S2814D), S2814D crossed with PLNKO mice (SDKO), and a previously validated human cardiac myocyte model. Restitution of cytosolic Ca²⁺ (Fura-2 AM) and L-type calcium current (ICaL; patch-clamp) was evaluated with a two-pulse (S₁/S₂) protocol. CRR and ICaL restitution increased as a function of the (S₁-S₂) coupling interval, following an exponential curve. When SR Ca²⁺ load was increased by increasing extracellular [Ca²⁺] from 2.0 to 4.0 mM, CRR and ICaL restitution were enhanced, suggesting that ICaL restitution may contribute to the faster CRR observed at 4.0 mM [Ca²⁺]. In contrast, ICaL restitution did not differ among the different mouse models. For a given SR Ca²⁺ load, CRR was accelerated in S2814D myocytes versus WT, but not in PLNKO and SDKO myocytes versus WT and S2814D, respectively. The model mimics all experimental data. Moreover, when the PLN ablation-induced decrease in RYR2 expression was corrected, the model revealed that CRR was accelerated in PLNKO and SDKO versus WT and S2814D myocytes, consistent with the enhanced velocity of refilling, SR [Ca²⁺] recovery, and CRR. We speculate that refilling rate might enhance CRR independently of SR Ca²⁺ load.
format Articulo
Articulo
author Cely Ortiz, Diana Catalina Alejandra
Felice, Juan Ignacio
Diaz Zegarra, Leandro Agustín
Valverde, Carlos Alfredo
Federico, Marilén
Palomeque, Julieta
Wehrens, Xander H. T.
Kranias, Evangelia G.
Aiello, Ernesto Alejandro
Lascano, Elena Catalina
Negroni, Jorge A.
Mattiazzi, Alicia Ramona
author_facet Cely Ortiz, Diana Catalina Alejandra
Felice, Juan Ignacio
Diaz Zegarra, Leandro Agustín
Valverde, Carlos Alfredo
Federico, Marilén
Palomeque, Julieta
Wehrens, Xander H. T.
Kranias, Evangelia G.
Aiello, Ernesto Alejandro
Lascano, Elena Catalina
Negroni, Jorge A.
Mattiazzi, Alicia Ramona
author_sort Cely Ortiz, Diana Catalina Alejandra
title Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
title_short Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
title_full Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
title_fullStr Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
title_full_unstemmed Determinants of Ca²⁺ release restitution: Insights from genetically altered animals and mathematical modeling
title_sort determinants of ca²⁺ release restitution: insights from genetically altered animals and mathematical modeling
publishDate 2020
url http://sedici.unlp.edu.ar/handle/10915/124703
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