Habitat complexity 7 macroinvertebrate 6 species richness 6 Crustacea 4 Ecosystem engineers 4 Macroinvertebrate assemblage 4 Orchestia gammarella 4 Patagonia 4 Polychaeta 4 Spartina 4 Thoracica 4 abundance 4 coastal zone 4 community response 4 dominance 4 ecosystem engineering 4 ecosystem management 4 habitat management 4 saltmarsh 4 spatial distribution 4 species diversity 4 Amphibia 2 Aves 2 Community structure 2 Diversity 2 Wetlands 2 aquatic plant 2 community structure 2 environmental factor 2 food availability 2